Spatial patterns and intra-specific competition of Juniper tree in different life stages using O- ring statistic in Layen Forests, Iran

Analyzing spatial patterns of individuals within life stages and spatial associations between different life stages is essential for understanding the spatial and temporal dynamics of populations. This research was conducted to study the spatial patterns and spatial association of Juniper trees (Juniperus polycarpos C. Koch) at different life stages in Juniper forests of Layen, Northeast of Iran. Data collection was done using four one- hectare square sample plots randomly selected, which fully mapped. Spatial patterns and spatial association of different life stages (juvenile, premature and mature; based on crown diameter) were then analyzed using O- ring univariate and bivariate statistics, respectively. Results showed that the general spatial patterns of the trees in total and in every life stage was clumped at least up to 25 m, while after that tends to random distribution at larger scales. Spatial association analyses revealed that there was no significant intra-specific competition among life stages namely, the spatial association was positive (attraction) or indifferent. It is concluded that clump patterns are due to heavy fruit and dispersal limitation of Juniper trees and harsh environmental condition of the study area.