جستجوی مقالات مرتبط با کلیدواژه "essential oil content" در نشریات گروه "باغبانی"

To observe the effect of plant growth-promoting rhizobacteria (PGPR) and salicylic acid under drought stress conditions on yarrow, an experiment was arranged as split-plot factorial based on a randomized complete block design with three replications in a research farm in Basht, Kohgiluyeh and Boyer-ahmad provice, during two years (2018 and 2019). Drought stress as the main factor in two levels (drought and non-drought stress), and secondary factors include foliar spraying with salicylic acid in two levels: control (no foliar spraying) and foliar spraying with salicylic acid (400 mg/L), and PGPR at four levels: control (no inoculation with bacteria), inoculation with Pseudomonas, inoculation with Azotobacterial and inoculation with both bacteria were assigned to the main and sub-plots, respectively. The results of a two-year combined variance analysis showed that effects of year, drought stress, bacteria, salicylic acid and drought stress × salicylic acid × bacteria on plant height, thousand seed weight, flower yield, biological yield, essential oil content, and essential oil yield were significant. The average flower yield in the first year was 731.45 kg/ha, which had a significant difference compared to the second year. The highest plant height (65.94 cm) was obtained from azotobacterial and salicylic acid treatment under non-drought stress conditions. Meanwhile, in both conditions (non-drought stress and drought stress), the maximum thousand seed weight (0.29 and 0.23 g), flower yield (1389 and 492 kg/ha), biological yield (8634 and 4106 kg/ha), harvest index (16.09 and 11.99 %), essential oil content (1.41 and 0.80 %) and essential oil yield (1.77 and 19.54 kg/ha) were obtained for bacteria and salicylic acid treatment. Although drought stress caused a significant decrease in all traits, especially flower yield, essential oil content and yield, the use of growth-promoting bacteria or salicylic acid, especially in the treatment of simultaneous application of two bacteria and salicylic acid, significantly reduced the damage of drought stress on flower yield, essential oil content and yield compared to the control.

To evaluate the effects of different irrigation levels and foliar application of nutrients, on dry matter yield and essential oil content and composition of peppermint (Mentha piperita L.), a field experiment was conducted as a split plot based on a randomized complete block design, with 12 treatments and three replications, at the faculty of Agriculture, University of Maragheh, Iran in 2021. The factors included irrigation as the main factor at three levels (irrigation after 20 (control), 40 (moderate stress), and 60% (severe stress) of the maximum allowable moisture depletion) and fertilizer source as the sub-factor at four levels (non-application of fertilizer (control), and foliar application of Macromix gold, Ecoquel micromix, and Vitalem forte). The results demonstrated that the highest (280.5 g.m-2) and lowest (113.7 g.m-2) total dry weight were obtained under normal irrigation conditions (control) with the application of Macromix gold and severe stress without fertilizer application, respectively. Total dry weight decreased by 22.7 and 40.3% under moderate and severe drought stress conditions, respectively compared to normal irrigation. Macromix gold, Vitalem forte, and Ecoquel micromix application enhanced the amount of total dry weight by 37.7, 23.8, and 17.1% compared to no fertilizer application, respectively. The highest content (2.1%) and essential oil yield (4.5 g.m-2) and also the highest amount of menthol (40.9%), menthone (28.1%), and cineole (6.2%) in the essential oil were observerd under moderate water stress conditions fertilized with Macromix gold. Based on the obtained results, the foliar application of Macromix gold could be recommended as an effective solution to enhance the quantity and quality of the peppermint essential oil under drought stress conditions.

Piriformospora indica is one of the growth-promoting microorganisms that can stimulate growth and increase plant tolerance in adverse environmental conditions. To evaluate the effects of P. indica inoculation on some vegetative, physiological, biochemical, and nutrient acquisition parameters and essential oil content of Origanum vulgare L. ssp. vulgare, a pot experiment was conducted in a completely randomized design with three replications. Treatments included control (without inoculation) and inoculation with P. indica. The results showed that inoculation with P. indica increased growth parameters, chlorophyll index, total phenols and total flavonoids content, essential oil content, and nutrients uptake of iron, potassium, and phosphorus singnificantly. The highest and lowest amounts for aerial parts fresh yield (3.92 and 3.06 g plant-1), aerial parts dry yield (1.12 and 0.78 g plant-1), chlorophyll index (47 and 42.23), total phenol (4.88 and 2.96 mg GAE g-1 FW), total flavonoids (0.52 and 0.23 mg QE g-1 FW), essential oil content (1.43 and 1.01%), phosphorus (0.41 and 0.35%), potassium (3.8 and 2.6%), and iron (219 and 180 mg kg-1) were obtained in the plants inoculated with fungus and non-inoculated plants, respectively. Inoculation with P. indica decreased the amount of major essential oil compounds such as carvacrol and thymol and increased the amount of ρ-cymene, carvacrol methyl ether, and γ-terpinene compared to the control. Overall, the findings of this study showed that inoculation with growth-promoting microorganisms can improve the morphological and phytochemical traits of oregano by improving nutrients uptake.

In the arid and semiarid regions, water stress mainly affects the growth, yield and secondary metabolit production of oregano. In order to determine the effect water stress on growth, physiological and phytochemicals characteristic of American oregano (Origanum vulgare L.), a greenhouse study based on a completely randomized design (CRD) with four treatments and three replications was performed. The plants, were subjected to four water stress conditions, including no water stress (field moisture capacity), mild (75 % FMC), moderate (55 % FMC) and sever water stress (35 % FMC). The results showed that the plant height, dry matter yield, the ratio of inflorescence weight/stem, chlorophyll index and relative water content were reduced under water stress. The highest essential oil content (1.15 %) was observed under mild water stress. Also, mild water stress significantly decreased the chlorophyll fluorescence parameters (Y(NO) and Fv) in compared with moderate and severe water stress. Analysis of the essential oil components demonstrated that in all treatements, sabinene, (Z)-β-ocimene, (E)-β-ocimene, (E)-caryophyllene, germacrene D and γ-terpinene were the major components. The highest amount of γ-terpinene (16.79 %) and (E)-caryophyllene (9.92 %) were observed in mild water stress which increased by 344.1 % and 14.28 % compared with control, respectively. Based on the obtained results, mild water stress significantly ameliorate the quality and quantity of oregano.

To study the effects of different nitrogen levels (0, 70, 140, and 210 mg l-1) on some growth, biochemical, and nutrient uptake (nitrogen, phosphorus, and potassium) characteristics and essential oil content in sage (Salvia officinalis L.) under hydroponic conditions, a pot experiment was conducted based on a completely randomized design with four replications in the research greenhouse of Ferdowsi University of Mashhad in 2019. The results of means comparison showed that with increasing the nitrogen level, the plant growth characteristics increased, so that the 210 mg l-1 N treatment decreased the stem diameter, number of branches, nodes, and leaves, length of internodes, length and leaf width, fresh and stem dry weight, fresh and aerial parts dry weight, and aerial parts dry weight to root dry weight ratio by 54.19, 72.59, 39.49, 82.52, 51.42, 72.20, 71.46, 95.17, 90.31, 95.82, 92.30, and 83.46%, respectively compared to the no N application. The application of 210 mg l-1 N increased the relative content of leaf water (RWC), proline, and essential oil content by 9.57, 266.6, and 14.87% and decreased the electrolyte leakage and malondialdehyde by 30.45 and 51.51%, respectively compared to the no N application. Also, the lowest N level decreased the K, P, and N content by 32.65, 85.13, and 56.89%, respectively compared to the highest N level. The results of this experiment showed that different N levels could have different effects on the growth and essential oil content of sage. Based on the present research findings, the application of 210 mg l-1 N could be recommended for the sage cultivation due to the improving effects on the studied traits of the plant.

In order to evaluate the integrative application of arbuscular mycorrhizal fungi and chemical fertilizers on the quality and quantity traits of Dragon's head (Lallemantia iberica), a field experiment was carried out as a randomized complete blocks design (RCBD) with eight treatments and three replications at the faculty of Agriculture, University of Maragheh during 2016 growing season. Treatments were included 100% chemical fertilizer (CF), Glomus mosseae (GM), Glomus intraradices (GI), GM+GI, 50% CF+GM, 50% CF+GI and 50% CF+GM+GI. The results demonstrated that the highest (71.83) and the lowest (54.47) chlorophyll index was achieved in the 50% CF+GM+GI and individual application of GI. In addition, the highest number of lateral branches was related to application of 50% CF+GM+GI, 50% CF+GM and GI. Furthermore, the highest grain yield (802.8 Kg ha-1), number of flower cycles per plant (63.6), number of seed per plant (519.3), number of seeds in the main stem (149.2), number of seeds per flower cycle (18), seed weight per plant (3.75 g), the grain-1000 weight (4.75 g) and total dry weight (426.3 g m-2) were achieved in the integrative application of 50% CF+GM+GI. By the way, the highest of EO content (0.5%) and EO yield (2.2 g m-2) were observed with application of 50% CF+GM+GI. The analysis of GC-FID and GC-MS showed that the linalool,menthone,geranial,menthyl acetate,geranyl acetate,valencene andcaryophyllene oxide were the major essential oil compounds. Generally, integrative application of 50% CF+GM+GI could improve the quality and quantity traits of dragon's head.

Ensuring the healthy and sustainable production of agricultural products, esp. medicinal plants, while preserving the environment, is an important issue that is becoming increasingly important every day. To investigate the effect of seaweed fertilizer foliar application on growth characteristics and quantitative and qualitative yield of hyssop (Hyssopus officinalis L.), an experiment was conducted in a randomized complete block design with four treatments of seaweed fertilizer (0, 2.5, 5 and 10 ml.l-1) and three replications in the research farm of faculty of agriculture, Tarbiat Modares University, Tehran province, Iran in 2017. Measured traits included plant height, leaf area, stem diameter, top flowering branch length, fresh and dry weight of the plant, chlorophyll content index, yield, essential oil content, and composition. The results showed that seaweed fertilizer had a significant effect on all studied traits except top flowering branch length and stem diameter, and the highest plant height (49.66 cm), fresh and dry weight of the plant (103.22 and 22.89 g, respectively), leaf area (0.77 cm2) and chlorophyll content (61.83) were obtained in 10 ml.l-1 seaweed fertilizer treatment without significant difference from 5 ml.l-1 treatment. Also, the 10 ml.l-1 treatment caused the highest percentage and yield of essential oil (0.58% and 2.12 g.m-2). Essential oil analysis showed that β-pinene, trans-pinocamphone, and cis-pinocamphone were the main compounds of essential oil in all treatments. The highest content of β-pinene (32.8%), cis-pinocamphone (42.5%), and trans-pinocamphone (19.6%) was observed in 5, 2.5 and 10 ml.l-1 seaweed fertilizer treatments, respectively, without significant difference from control treatment. Based on the results of this study, the application of seaweed fertilizer can improve the growth characteristics and quantitative and qualitative yield of hyssop.

Drought stress (water scarcity) is one of the most limiting environmental factors in plant growth and production. Sodium nitroprusside (SNP) as a nitric oxide doner compound (NO) plays an important role in reducing the oxidative stress caused by abiotic stresses. In order to investigate the effect of SNP foliar application on some biochemical characteristics, growth and yield of Echinacea purpurea Moench L. under drought conditions, an experiment was conducted in a split plot design in a completely randomized block design with three replications at the Research Institute of Agriculture, Institute of Forest and Rangelands, Karaj, Iran, in 2015-2016. The experimental treatments included SNP factor at four levels (0, 100, 200 and 400 µM) and drought stress factor at three levels [30% field capacity (severe), 60% field capacity (moderate), 90% field capacity (control)]. At full flowering stage, plant height, number of lateral branches, leaf and flower number, dry weight of shoots and percentage and yield of shoot essential oil were measured. Also, leaves sampling was performed to measure relative water content (RWC), chlorophyll content, the activity of superoxide dismutase (SOD) and peroxidase (POD) enzymes, malondialdehyde (MDA), total phenol and proline contents. Results showed that the hieghest amount of morphological triats and shoot dry weight were related to 200 µM SNP concentration under non drought stress (control) condition. The highest amount of RWC was obtained (97.9%) in 90% moisture treatment (control) and 200 µM of SNP. Also, higher concentrations of SNP (200 and 400 µM) under moderate stress (60% of crop capacity), could increase the plant drought stress tolerance by increasing the SOD and POD enzymes activity as well as reducing MDA content. The highest amount of proline (16.7 µmol/g fr wt) and total phenol (127.7 mg GAE/g dry matter) were also related to 400 µM SNP treatment. In general, SNP foliar application especially at 200 µM concentration under drought stress conditions could modify drought stress damage, and increase active ingredient of the plant.

In order to evaluate the effects of foliar application of osmolytes on the quantitative and qualitative yield of German chamomile (Chamomilla recutita L.) in drought stress conditions, a split-plot experiment was conducted based on randomized complete block design with three replications at Research Center of Medicinal Plants, Shahed University. The treatments included irrigation as the main factor in three levels [(irrigation after 50 (without stress), 100 (moderate stress) and 150 mm evaporation from the class A pan (severe stress)] and foliar application of osmolytes as sub-plots in seven levels [(without Spraying (NS), distilled water (W), methyl jasmonate (MJ= 75mM), salicylic acid (SA= 2mM), humic acid (HA= 2.5lit/1000), glycine betaine (GB= 5mM), and g- aminobutyric acid (GABA= 50mM). The electrical conductivity of the irrigation water was 4.8 dS/m. Mean comparisons showed that GABA combined with severe stress had the highest essential oil yield (3.7 kg ha-1). In plants treated with GABA under severe stress, essential oil harvest index was increased by 60 and 58%, respectively, compared to W and NS treatments. The highest and lowest capitol harvest index with 70.81% and 23.28% were respectively obtained from GB combined without stress and W with severe stress. SA, GB, and GABA increased the capitol yield in severe stress compared with the moderate stress. Severe stress caused a significant increase in essential oil content from 0.66 to 1.13%, a 35% reduction in seed harvest index compared to control treatment, and a significant reduction in biomass yield rather than moderate stress. MJ and SA increased seed yield compared with NS in moderate stress.

In order to investigate the effect of drought stress on grain and essential oil yield, antioxidant properties of essential oil, and accumulation of conformal osmolites in the leaves of two Nigella sativa L., this research was conducted at the research farm of Islamic Azad University, Shoushtar Branch during 2016-2017. The experiment was a split plot based on randomized complete block design with four replications. The main plot included four levels of irrigation including optimal irrigation, mild, moderate and severe stress (based on irrigation under 90, 80, 70 and 60% soil field capacity, respectively), and ecotypes of Kashmar and Shahreza were considered as subplots. Results indicated that the grain yield of Kashmar ecotype was reduced 2.4%, 20.2% and 40.1% under mild, moderate and sever stress compared to optimum conditions, respectively. The grain yield of Shahreza ecotype was reduced 6.2%, 38.1% and 63%, respectively. Reduction of grain yield under severe stress conditions was due to a significant decrease in the number of capsules and seeds per capsule. Essential oil yield of Kashmar and Shahreza ecotypes decreased by 18% and 54% in severe drought stress conditions, respectively. Kashmar ecotype had higher levels of carbohydrate and proline than those of Shahraza under stress conditions. This property caused the Kashmar ecotype to have more relative water content and lower slope change of grain yield and essential oil yield as compared with Shahreza. In normal conditions and severe drought stress, 85% and 93% of essential oil components of Kashmar and 82% and 93% essential oil components of Shahreza ecotype were identified, respectively. Drought stress increased the percentage of essential oil in both ecotypes and the essential oil of Kashmar contained more antioxidant. In general, under mild stress conditions, the highest essential oil yield was obtained due to an increase in essential oil content and no significant reduction in grain yield. Thus, it seems that the application of mild controlled stress improves the quality of Nigella sativa without affecting its quantitative performance.

Today, in a sustainable agriculture system, application of the fertilizers which are nature friendly and suitable for plants is essential. This becomes more important when dealing with medicinal plants. Therefore, a factorial experiment in a randomized complete block design with three replications was set up in the 2011-2013 on Nigella sativa. Treatments were Nano fertilizer (Farmks®) (at 2 levels 0, 1 ml/l) and humic acid (at 4 levels zero, 1, 3, 6 ml/l) which were applied at three time of plant growth. Results showed that different levels of humic acid imposed a significant effect on plant height, leaf area index, dry weight, number of branches, and number of follicle per plant, number of seeds per follicle, seed weight, seed yield, biological yield harvest index, essential oil content and yield. The highest yield was in levels of 3, 6 ml/l of humic acid treatments, respectively. Nano fertilizer (Farmks®) application significantly increased the yield and yield components of Nigella sativa. Combined treatment at various levels had significant effect on dry weight, seed weight, seed yield, biological yield and harvest index respectively but had no significant effect on other traits. According to these results, we can conclude that the humic acid and Nano fertilizer (Farmks®) application could increase yield, yield components, essential oil contentand and yield.also, combination treatment (humic acid and Nano fertilizer) ware more effective in increase of essential oil content and yield. Therefore, the application of this kind of fertilizers can be advisable in Nigella sativa.

Foliar application of micronutrients is a useful technique for rapid absorption of minerals in the plants and avoids soil contamination. To evaluate the effect of foliar application of micronutrients (iron and zinc) on yield, yield components, and essential oil of fennel (Foeniculum vulgare Mill.), an experiment was conducted as factorial based on randomized complete blocks design with three replications at the Faculty of Agriculture, University of Zanjan during the growing season of 2012. In this experiment, the effect of foliar application of Fe (using ferrous sulphate) at four concentrations (0, 1, 2 and 3 gr.L-1) and foliar application of Zn (using zinc sulphate) at three concentrations (0, 2 and 4 gr.L-1) were investigated. Results showed that the majority of measured traits including number of umbels per plant, 1000-seed weight, grain yield, essential oil content and essential oil yield were significantly affected (p≤ 0.01) by combined application of iron and zinc. Therefore, the highest grain yield (2637.96 kg ha-1) was obtained in foliar application of Fe + Zn at a concentration of 0.2%. In addition, the highest essential oil content (2.88%) and essential oil yield (75.71 kg ha-1) were obtained in combined application of Fe 0.3% and Zn 0.2% and the lowest content of mentioned traits was obtained in control.

To study the effects of seed priming and foliar application of salicylic acid (SA) on yield and essence of fennel (Foeniculum vulgar Mill.) under drought stress condition, a field experiment was conducted at College of Agriculture, University of Zanjan in the spring of 2012. The experiment was conducted as a split plot based on complete randomized block design with three replications. In this study, drought stress at reproductive stage with control (full irrigation) was assigned to the main plots and different treatments of seed priming and foliar application with SA including dry seeds (without priming), seeds priming with SA at different concentrations (0, 800 and 1600μmol) and foliar application with SA at different concentrations (800 and 1600μmol) were assigned to the subplots. Results showed that the majority of measured traits including number of umbrella in plant, number of seeds in umbrella, seed thousand weight, harvest index, biomass, grain yield, and essential oil yield significantly decreased under drought stress condition and the essential oil content increased vice versa. Therefore, the highest seed (2925.6kg/ha) and essential oil yield (52.88kg/ha) were obtained in control, and the lowest amount of mentioned traits was obtained in drought stress condition. The results also showed that the effect of seed priming and foliar application of SA on the majority of measured traits was significant. There were no differences about seed yield and essential oil yield between foliar application and priming with SA at a concentration of 1600 μmol; however, the highest seed yield (2579.60kg/ha) and essential oil yield (63.15kg/ha) were obtained in foliar application with SA at a concentration of 1600μmol, and the lowest amounts of mentioned traits were obtained in dry seeds (without priming).

Sweet majoram (Majorana hortensis L.) is one of the native Iranian medicinal plants، which due to its aromatic secondary metabolites is used in traditional medicine. In production of medicinal plants، application of organic inputs has especial role in sustainable agriculture. Organic inputs، such as manures and compost، other than providing the necessary elements for plant growth، could improve the physiochemical structure of soil. This research was conducted to evaluate the growth traits، quantitative and qualitative yield، and constituents of essential oil of sweet majoram under the application of compost extract and inoculation of biofertilizers. The experiment was performed as split plot in time with three replications at Research Greenhouse of Ferdowsi University، Mashhad، Iran، in 2012. Main plots were 16 fertilizer combinations (Azotobacter (Azotobacter paspali)، (B1)، Azospirillum (Azospirillum brasilense) (B2)، mycorrhiza (Glomus intraradaices) (B3)، 10% compost extract (C1)، 20% compost extract (C2)، 30% compost extract (C3)، B1C1، B1C2، B1C3، B2C1، B2C2، B2C3، B3C1، B3C2، B3C3 and control) and sub- plots were three cutting stages during the growing season. Results showed that height of main stem، dry and fresh weight of sweet majoram was significantly affected by single and combined application of biofertilizer inoculation and compost extract at the first cutting and were higher than the second and third cuttings. Also، the most significant increase in height of main stem، fresh and dry weight and essential oil content of sweet majoram (43. 3 cm، 46. 07 g/m2، 27. 11 g/m2 and 0. 89%، respectively) was observed in C1B3 treatmeent. Constituents of essential oil were significantly affected by single and combined applications of biofertilizer inoculation and compost extract. Among the constituents، terpinen-4-ol (18. 01%) and cis-sabinene hydrate (15. 4%) allocated the highest and ledene (15. 4%) and camphene (0. 01%) the lowest amount.

This research was aimed to investigate the yield and quality response of two endemic chamomile (Matricaria chamomilla L.) ecotypes from Bushehr and a commercial ecotype from Esfahan to biofertilizers، a field experiment was conducted during two growing seasons of 2008 and 2009 at the research farm of Persian Gulf University (Boushehr campus). The experiment was carried out in a randomized complete blocks design in a factorial arrangement with three replications. Treatments consisted of chamomile ecotypes، mycorrhisal inoculationfection (with and without) and amount of micro-biophosphate fertilizer (0، 30 and 60kg. ha-1). Morphological traits and flower yield were evaluated from the flowering period onwards and trhen the essential oil concentration، Chamazulene and Apigenine 7-glycoside percentage were measured. Also، after final harvest، nitrogen، phosphorous and potassium content of chamomile plant and soil were investigated. Result showed that except of the main effect of ecotypes، the other main and interaction effects on the measured traits were insignificant. The flower dry weight of Bushehr ecotypes (1 and 2) was %34 more than that of Esfahan ecotype in 1st year. However، in 2nd year، Boushehr2 had the highest dry flower yield (1132. 66 kgha-1) followed by Boushehr1 and Esfahan ecotypes with 12. 4 and 48. 8 percent loss، respectively. In both years of experiment، Esfahan ecotype produced more chamazulene in essential oil and Boushehr ecotypes were superior treatments for Apigenine 7-glycoside in dried flower (p≤0. 05). Although our finding revealed no significant effect of biofertilizers on all measured traits، a dry flower yield of 800-1000 kgha-1، 3 kg ha-1 essential oil، high percentage of chamazulene in essential oil (15-16% for Esfahan and 5-7% for Boushehr ecotypes)، and considerable amount of Apigenine 7-glycoside in dried flower (10-12. 7 kgha-1) demonstrated the high potential yield of chamomile using biofertilizers under field condition. Therefore، since chamomile showed a proper and vast ecological adaptation to the cultural conditions in Boushehr region، it could be introduced to the low input agricultural systems as a reliable part of local crop rotations.

Drying is one of the oldest methods for preservation of agricultural products after harvest. Basil (Ocimum basilicum L.) belongs to Lamiaceae family which is used as a medicinal and spice species. This experiment was carried out to determine the effect of different drying methods on drying time, essential oil content and composition of Basil. The experiment was conducted in a randomized complete blocks design with three replications and treatments were included two temperature levels (50 and 70 °C), six microwave powers (100, 180, 300, 450, 600 and 900 w), and two drying methods (sun and shade drying). The drying process was continued until the mass of the sample was reduced to a moisture content of about 0.10 on a dry basis or 10 % on a wet basis. Results indicated minimum and maximum drying times (4.1 minutes and 48 hours) were obtained at 900 w microwave powers and shade drying, respectively. Maximum and minimum essential oil content (1.3 and 0.3 %) were obtained by shade drying and in 450, 600 and 900 w, respectively. Microwave powers had negative effect on the main compositions of essential oil (geranial and methyl chavicol) but shade drying preserved them. According to the results, shade drying was identified as a suitable method for Basil, because essential oil content and composition were considerably maintained.

In order to evaluate the effects of water salinity on ajowan (Carum copticum L.), a completely randomized design was conducted in Esfahan province in 2007. Eight water salinity treatments (0.3-21 dS/m) with 3 replications were applied in the field. The seeds (at ripening) and foliage (initial flowering) of ajowan were harvested. After drying the plant materials in shade, their essential oils were obtained by hydro-distillation. The oils were analyzed by capillary gas chromatography, using flame ionization and mass spectrometric detection. According to results seed and foliage oils percentage were 3.5-4.4 and 0.25-0.35 cc/100 g, respectively. Maximum seed and foliage oils yield were 8.3 and 2.4 cc/m2 and minimum were 3.5 and 1.5 cc/m2 in control and in 21 dS/m, respectively. Maximum biological and seed yields were 202.7 and 1367.6 and minimum were 96.7 and 739.5 gr/m2 in control and 21dS/m, respectively. The most important constituents of essential oil were: thymol (55.5% and 56.2%), γ-terpinene (22.5% and 26.9%), ρ-cymene (14.2% and 11.2%), limonene (1.9% and 0.5%) and carvacrol (0.3% and 1.4%), respectively.